The top of molar cusps is relatively bunoselenomorph
He has an initial and straight postprotocrista labially dependent. The latest preprotocrista, apparently upright, meets a properly-setup parastyle. The metaconule are a bit reduced which can be huge with the M3. The fresh premetaconulecrista as well as the fossae are simple rather than neocrista. The latest posterior fossa are tilted, particularly on M3. Away from straight toward M1, brand new postmetaconulecrista will get sigmoidal toward M3, skipping brand new metacone. The fresh labial cusps have been in a comparable axis with apparently upright cristae. New paracone labial rib are salient, however, instead anterior groove. There’s absolutely no metacone labial rib. The fresh new para poder-, meso-, and you may metastyles is actually outstanding. The latest lingual cingulum is really poor in order to absent including the entostyle.
The brand new registered build of your last lower premolar is common getting derived base pecoran and you will crown pecoran during the having a properly-arranged mesolingual conid and you can posterior cristid , . More over, the current presence of an enamel outlet for the p1 excludes an attribution so you’re able to Prodremotherium or Mosaicomeryx . Even if the white teeth are not while the bunoselenodont as with Amphitragulus elegans, the fresh new called specimens have quite primitive enjoys, such as for instance a very quicker metastylid, a decreased top, a tiny postentocristid, and even though laterally compressed, bulged lingual cuspids into the lower molars. Each one of these attributes exclude an affiliation in order to both Dremotherium otherwise Bedenomeryx , . Additionally, the fresh new very small-size out-of “Amphitragulus” quercyi is a pinpointing ability to the newest Oligocene ruminants away from European countries , , , .
Dremotherium cf. guthi away from Gaimersheim1: step one kept maxillary bones for the enamel line M1-M3 (SNSB-BSPG 1952II4855). from Mailhat: dos remaining P4-M2 (MNHN LIM154). Babameryx engesseri off Gaimersheim1: step three leftover fragmentary mandible with m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you may (c) occlusal opinions. The shape bar was 2 cm.
Holotype introduced specimens recommendations
Holotype. Jehenne (1987) enjoys designated the latest sample IPHEP LM.1969.MA.40 since holotype. This might be a limited best maxillary limbs that have P3-M3 (, Fig. 2), of Los angeles Milloque (MP29, France).
Referred specimens. Mailhat MNHN LIM154 kept P4-M2 (so it really works, Fig. six.2); Gaimersheim1 SNSB-BSPG 1952II4855 kept maxillary which have M1-M3 (this performs, Fig. six.1); SNSB-BSPG 1952II4870 proper m2.
Breakdown
Lower permanent dentition. The low molar is very defectively kept. However, that it huge ruminant features a fully selenomorph trigonid. This new cristid is elongated while the cuspids is actually higher to the enamel quite pleated. The fresh metastylid was ripped.
Top permanent dentition. The newest P4 is actually badly kept. All lingual area are shed. The new anterolabial cone and also a failure labial rib. The new anterior stylid was broken. The new posterior you’re brief however, establish. Zero neocrista obstructs the new fossa. This new quadratic upper molars from inside the occlusal look at sustain a well-create metaconule some smaller compared to the newest protocone. New cusps are selenomorph. The fresh new preprotocrista together with postmetaconulecrista is actually labially mainly based and you can slightly 
straight. The fresh postprotocrista are much time, very rounded, and you may labially oriented. It could be totally separated (SNSB-BSPG 1952II4855) otherwise fused (MNHN LIM154) to your premetaconulecrista. The second finishes at the axis of your own mesostyle. The new rear basin normally keeps a little metaconule bend on the M2, big to your M3. The fresh paracone, having a paracone rib, and also the metacone, labially flat and you can concave, is actually aimed. The fresh mesostyle and you will parastyle are very well install. They function round brief articles. The fresh mesostyle is actually huge away from M1 so you can M3. The brand new entostyle was weakened otherwise absent. You will find none a beneficial lingual cingulum nearby brand new protocone neither an effective metastyle creating a well-create cuspid. Although not, small prior and you will rear cingula exist.
Talk
Which ruminant is relatively advanced in becoming really-selenodont and achieving no lingual cingula into the higher molars, usually found in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you can Babameryx , , , [61–63], . As opposed to Bachitherium, which types does not have an prior groove with the huge labial paracone rib , , . The analyzed fossils be a little more just like the advanced base Pecora and to new primitive top Pecora (such Amphitragulus, Dremotherium, Bedenomeryx). The fresh new enamel is actually a bit pleated, whereas it is generally simple inside the Prodremotherium. Also, the dimensions of this type of pearly whites is significantly larger than from inside the Prodremotherium regarding old Quercy selections and Pech Desse . One’s teeth are more selenomorph compared to “Amphitragulus” and you can Bedenomeryx , , as well as the parastyle isn’t that anteriorly projected than simply noticed towards “Amphitragulus” feningrei , , . Studying the neodiagnosis recommended because of the Mennecart , these fossils were all of the proving attributes of Dremotherium. The brand new currently laid out varieties inside European countries is D. feignouxi (sort of types) and you will D. guthi , , . The latter was slightly smaller compared to the kind variety , just as the fossils explained here. D. feignouxi just taken place during the Agenian (MN1-MN2; ), while D. guthi existed inside newest Oligocene (MP28-MP30; ). However, inside lack of extra well-kept material, a definitive assignation with the varieties isn’t feasible.